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In addition, MLC phosphorylation or activation of other kinases triggers the opening of TJ proteins (including ZO-1 and claudin-1), which can be reversed by adding an inhibitor of MLC phosphorylation [57]
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This was confirmed in an additional cell line and with depletion of KG or suppression of KGDDs using knockdown/inhibition of IDH1, glutamine starvation, or succinate supplementation ( Fig
