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To visualize small-molecule metabolites localized within TLS, we used matrix-assisted laser desorption/ionization time of flight imaging mass spectrometry and found significant accumulation of glutathione (reduced form: glutathione [GSH], oxidized form: glutathione disulfide [GSSG]) within TLS (Figure 1C)

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Recent findings have demonstrated that glutamine-addicted breast cancer cells exhibit a capacity to adapt to chronic glutamine starvation or GLS inhibition through AMPK-mediated upregulation of the serine synthesis pathway